The four remaining mammals are Marsupials, an order of the class Mammalia, which is very characteristic of the Australian fauna;and these are probably true natives of the Moluccas, since they are either of peculiar species, or if found elsewhere are natives only of New Guinea or North Australia. The first is the small flying opossum, Belideus ariel, a beautiful little animal, exactly line a small flying squirrel in appearance, but belonging to the marsupial order. The other three are species of the curious genus Cuscus, which is peculiar to the Austro-Malayan region. These are opossum-like animals, with a long prehensile tail, of which the terminal half is generally bare. They have small heads, large eyes, and a dense covering of woolly fur, which is often pure white with irregular black spots or blotches, or sometimes ashy brown with or without white spots. They live in trees, feeding upon the leaves, of which they devour large quantities, they move about slowly, and are difficult to kill, owing to the thickness of their fur, and their tenacity of life.
A heavy charge of shot will often lodge in the slain and do them no harm, and even breaking the spine or piercing the brain will not kill them for some hours. The natives everywhere eat their flesh, and as their motions are so slow, easily catch them by climbing; so that it is wonderful they have not been exterminated. It may be, however, that their dense woolly fur protects them from birds of prey, and the islands they live in are too thinly inhabited for man to be able to exterminate them.
The figure represents Cuscus ornatus, a new species discovered by me in Batchian, and which also inhabits Ternate. It is peculiar to the Moluccas, while the two other species which inhabit Ceram are found also in New Guinea and Waigiou.
In place of the excessive poverty of mammals which characterises the Moluccas, we have a very rich display of the feathered tribes. The number of species of birds at present known from the various islands of the Molluccan group is 265, but of these only 70 belong to the usually abundant tribes of the waders and swimmers, indicating that these are very imperfectly known. As they are also pre-eminently wanderers, and are thus little fitted for illustrating the geographical distribution of life in a limited area, we will here leave them out of consideration and confine our attention only to the 195 land birds.
When we consider that all Europe, with its varied climate and vegetation, with every mile of its surface explored, and with the immense extent of temperate Asia and Africa, which serve as storehouses, from which it is continually recruited, only supports 25l species of land birds as residents or regular immigrants, we must look upon the numbers already procured in the small and comparatively unknown islands of the Moluccas as indicating a fauna of fully average richness in this department.
But when we come to examine the family groups which go to make up this number, we find the most curious deficiencies in some, balanced by equally striking redundancy in other. Thus if we compare the birds of the Moluccas with those of India, as given in Mr. Jerdon's work, we find that the three groups of the parrots, kingfishers, and pigeons, form nearly _one-third_ of the whole land-birds in the former, while they amount to only _one-twentieth_ in the latter country. On the other hand, such wide-spread groups as the thrushes, warblers, and finches, which in India form nearly _one-third_ of all the land-birds, dwindle down in the Moluccas to _one-fourteenth._The reason of these peculiarities appears to be, that the Moluccan fauna has been almost entirely derived from that of New Guinea, in which country the same deficiency and the same luxuriance is to be observed. Out of the seventy-eight genera in which the Moluccan land-birds may be classed, no less than seventy are characteristic of Yew Guinea, while only six belong specially to the Indo-Malay islands. But this close resemblance to New Guinea genera does not extend to the species, for no less than 140 out of the 195 land-birds are peculiar to the Moluccan islands, while 32 are found also in New Guinea, and 15 in the Indo-Malay islands. These facts teach us, that though the birds of this group have evidently been derived mainly from New Guinea, yet the immigration has not been a recent one, since there has been time for the greater portion of the species to have become changed. We find, also, that many very characteristic New Guinea forms lave not entered the Moluccas at all, while others found in Ceram and Gilolo do not extend so far west as Bouru. Considering, further, the absence of most of the New Guinea mammals from the Moluccas, we are led to the conclusion that these islands are not fragments which have been separated from New Guinea, but form a distinct insular region, which has been upheaved independently at a rather remote epoch, and during all the mutations it has undergone has been constantly receiving immigrants from that great and productive island. The considerable length of time the Moluccas have remained isolated is further indicated by the occurrence of two peculiar genera of birds, Semioptera and Lycocorax, which are found nowhere else.