He hunts at night. This is as nearly invariable a rule as can be formulated in regard to lions. Yet once, and perhaps twice, I saw lionesses stalking through tall grass as early as three o'clock in the afternoon. This eagerness may, or may not, have had to do with the possession of hungry cubs. The lion's customary harmlessness in the daytime is best evidenced, however, by the comparative indifference of the game to his presence then. From a hill we watched three of these beasts wandering leisurely across the plains below. A herd of kongonis feeding directly in their path, merely moved aside right and left, quite deliberately, to leave a passage fifty yards or so wide, but otherwise paid not the slightest attention. I have several times seen this incident, or a modification of it. And yet, conversely, on a number of occasions we have received our first intimation of the presence of lions by the wild stampeding of the game away from a certain spot.
However, the most of his hunting is done by dark. Between the hours of sundown and nine o'clock he and his comrades may be heard uttering the deep coughing grunt typical of this time of night. These curious, short, far-sounding calls may be mere evidences of intention, or they may be a sort of signal by means of which the various hunters keep in touch. After a little they cease. Then one is quite likely to hear the petulant, alarmed barking of zebra, or to feel the vibrations of many hoofs. There is a sense of hurried, flurried uneasiness abroad on the veldt.
The lion generally springs on his prey from behind or a little off the quarter. By the impetus his own weight he hurls his victim forward, doubling its head under, and very neatly breaking its neck. I have never seen this done, but the process has been well observed and attested; and certainly, of the many hundreds of lion kills I have taken the pains to inspect, the majority had had their necks broken. Sometimes, but apparently more rarely, the lion kills its prey by a bite in the back of the neck. I have seen zebra killed in this fashion, but never any of the buck. It may be possible that the lack of horns makes it more difficult to break a zebra's neck because of the corresponding lack of leverage when its head hits the ground sidewise; the instances Ihave noted may have been those in which the lion's spring landed too far back to throw the victim properly; or perhaps they were merely examples of the great variability in the habits of felis leo.
Once the kill is made, the lion disembowels the beast very neatly indeed, and drags the entrails a few feet out of the way. He then eats what he wants, and, curiously enough, seems often to be very fond of the skin. In fact, lacking other evidence, it is occasionally possible to identify a kill as being that of a lion by noticing whether any considerable portion of the hide has been devoured. After eating he drinks. Then he is likely to do one of two things: either he returns to cover near the carcass and lies down, or he wanders slowly and with satisfaction toward his happy home. In the latter case the hyenas, jackals, and carrion birds seize their chance. The astute hunter can often diagnose the case by the general actions and demeanour of these camp followers. A half dozen sour and disgusted looking hyenas seated on their haunches at scattered intervals, and treefuls of mournfully humpbacked vultures sunk in sadness, indicate that the lion has decided to save the rest of his zebra until to-morrow and is not far away. On the other hand, a grand flapping, snarling Kilkenny-fair of an aggregation swirling about one spot in the grass means that the principal actor has gone home.
It is ordinarily useless to expect to see the lion actually on his prey. The feeding is done before dawn, after which the lion enjoys stretching out in the open until the sun is well up, and then retiring to the nearest available cover. Still, at the risk of seeming to be perpetually qualifying, I must instance finding three lions actually on the stale carcass of a waterbuck at eleven o'clock in the morning of a piping hot day! In an undisturbed country, or one not much hunted, the early morning hours up to say nine o'clock are quite likely to show you lions sauntering leisurely across the open plains toward their lairs.
They go a little, stop a little, yawn, sit down a while, and gradually work their way home. At those times you come upon them unexpectedly face to face, or, seeing them from afar, ride them down in a glorious gallop. Where the country has been much hunted, however, the lion learns to abandon his kill and seek shelter before daylight, and is almost never seen abroad. Then one must depend on happening upon him in his cover.
In the actual hunting of his game the lion is apparently very clever. He understands the value of cooperation. Two or more will manoeuvre very skilfully to give a third the chance to make an effective spring; whereupon the three will share the kill. In a rough country, or one otherwise favourable to the method, a pack of lions will often deliberately drive game into narrow ravines or cul de sacs where the killers are waiting.
At such times the man favoured by the chance of an encampment within five miles or so can hear a lion's roar.
Otherwise I doubt if he is apt often to get the full-voiced, genuine article. The peculiar questioning cough of early evening is resonant and deep in vibration, but it is a call rather than a roar. No lion is fool enough to make a noise when he is stalking.
Then afterward, when full fed, individuals may open up a few times, but only a few times, in sheer satisfaction, apparently, at being well fed. The menagerie row at feeding time, formidable as it sounds within the echoing walls, is only a mild and gentle hint. But when seven or eight lions roar merely to see how much noise they can make, as when driving game, or trying to stampede your oxen on a wagon trip, the effect is something tremendous.
The very substance of the ground vibrates; the air shakes. I can only compare it to the effect of a very large deep organ in a very small church. There is something genuinely awe-inspiring about it; and when the repeated volleys rumble into silence, one can imagine the veldt crouched in a rigid terror that shall endure.