If, however, Zadig was right in concluding from the likeness of the hoof-prints which he observed to be a horse's that the creature which made them had a tail like that of a horse, Cuvier, seeing that the teeth and jaw of his fossil were just like those of an opossum, had the same right to conclude that the pelvis would also be like an opossum's; and so strong was his conviction that this retrospective prophecy, about an animal which he had never seen before, and which had been dead and buried for millions of years, would be verified, that he went to work upon the slab which contained the pelvis in confident expectation of finding and laying bare the "marsupial bones," to the satisfaction of some persons whom he had invited to witness their disinterment. As he says:--"Cette operation se fit en presence de quelques personnes a qui j'en avais annonce d'avance le resultat, dans l'intention de leur prouver par le fait la justice de nos theories zoologiques; puisque le vrai cachet d'une theorie est sans contredit la faculte qu'elle donne de prevoir les phenomenes."
In the "Ossemens Fossiles" Cuvier leaves his paper just as it first appeared in the "Annales du Museum," as "a curious monument of the force of zoological laws and of the use which may be made of them."
Zoological laws truly, but not physiological laws. If one sees a live dog's head, it is extremely probable that a dog's tail is not far off, though nobody can say why that sort of head and that sort of tail go together; what physiological connection there is between the two. So, in the case of the Montmartre fossil, Cuvier, finding a thorough opossum's head, concluded that the pelvis also would be like an opossum's. But, most assuredly, the most advanced physiologist of the present day could throw no light on the question why these are associated, nor could pretend to affirm that the existence of the one is necessarily connected with that of the other. In fact, had it so happened that the pelvis of the fossil had been originally exposed, while the head lay hidden, the presence of the "marsupial bones," though very like an opossum's, would by no means have warranted the prediction that the skull would turn out to be that of the opossum. It might just as well have been like that of some other marsupial; or even like that of the totally different group of Monotremes, of which the only living representatives are the Echidna and the Ornithorhynchus.
For all practical purposes, however, the empirical laws of co-ordination of structures, which are embodied in the generalisations of morphology, may be confidently trusted, if employed with due caution, to lead to a just interpretation of fossil remains; or, in other words, we may look for the verification of the retrospective prophecies which are based upon them.
And if this be the case, the late advances which have been made in palaeontological discovery open out a new field for such prophecies. For it has been ascertained with respect to many groups of animals, that, as we trace them back in time, their ancestors gradually cease to exhibit those special modifications which at present characterise the type, and more nearly embody the general plan of the group to which they belong.
Thus, in the well-known case of the horse, the toes which are suppressed in the living horse are found to be more and more complete in the older members of the group, until, at the bottom of the Tertiary series of America, we find an equine animal which has four toes in front and three behind. No remains of the horse tribe are at present known from any Mesozoic deposit.
Yet who can doubt that, whenever a sufficiently extensive series of lacustrine and fluviatile beds of that age becomes known, the lineage which has been traced thus far will be continued by equine quadrupeds with an increasing number of digits, until the horse type merges in the five-toed form towards which these gradations point?
But the argument which holds good for the horse, holds good, not only for all mammals, but for the whole animal world. And as the study of the pedigrees, or lines of evolution, to which, at present, we have access, brings to light, as it assuredly will do, the laws of that process, we shall be able to reason from the facts with which the geological record furnishes us to those which have hitherto remained, and many of which, perhaps, may for ever remain, hidden. The same method of reasoning which enables us, when furnished with a fragment of an extinct animal, to prophesy the character which the whole organism exhibited, will, sooner or later, enable us, when we know a few of the later terms of a genealogical series, to predict the nature of the earlier terms.
In no very distant future, the method of Zadig, applied to a greater body of facts than the present generation is fortunate enough to handle, will enable the biologist to reconstruct the scheme of life from its beginning, and to speak as confidently of the character of long extinct beings, no trace of which has been preserved, as Zadig did of the queen's spaniel and the king's horse. Let us hope that they may be better rewarded for their toil and their sagacity than was the Babylonian philosopher; for perhaps, by that time, the magi also may be reckoned among the members of a forgotten Fauna, extinguished in the struggle for existence against their great rival, common sense.