In every arduous enterprise it is pleasanter to look back at difficulties overcome than forward to those which still seem insurmountable, but in the next stage there is nothing to be gained by disguising the fact that the attributes of living things are not what we used to suppose. If they are more complex in the sense that the properties they display are throughout so regular (I have in view, for example, the marvellous and specific phenomena of regeneration, and those discovered by the students of "Entwicklungsmechanik". The circumstances of its occurrence here preclude any suggestion that this regularity has been brought about by the workings of Selection. The attempts thus to represent the phenomena have resulted in mere parodies of scientific reasoning.) that the Selection of minute random variations is an unacceptable account of the origin of their diversity, yet by virtue of that very regularity the problem is limited in scope and thus simplified.
To begin with, we must relegate Selection to its proper place. Selection permits the viable to continue and decides that the non-viable shall perish; just as the temperature of our atmosphere decides that no liquid carbon shall be found on the face of the earth: but we do not suppose that the form of the diamond has been gradually achieved by a process of Selection. So again, as the course of descent branches in the successive generations, Selection determines along which branch Evolution shall proceed, but it does not decide what novelties that branch shall bring forth. "La Nature contient le fonds de toutes ces varietes, mais le hazard ou l'art les mettent en oeuvre," as Maupertuis most truly said.
Not till knowledge of the genetic properties of organisms has attained to far greater completeness can evolutionary speculations have more than a suggestive value. By genetic experiment, cytology and physiological chemistry aiding, we may hope to acquire such knowledge. In 1872 Nathusius wrote ("Vortrage uber Viehzucht und Rassenerkenntniss", page 120, Berlin, 1872.): "Das Gesetz der Vererbung ist noch nicht erkannt; der Apfel ist noch nicht vom Baum der Erkenntniss gefallen, welcher, der Sage nach, Newton auf den rechten Weg zur Ergrundung der Gravitationsgesetze fuhrte."We cannot pretend that the words are not still true, but in Mendelian analysis the seeds of that apple-tree at last are sown.
If we were asked what discovery would do most to forward our inquiry, what one bit of knowledge would more than any other illuminate the problem, Ithink we may give the answer without hesitation. The greatest advance that we can foresee will be made when it is found possible to connect the geometrical phenomena of development with the chemical. The geometrical symmetry of living things is the key to a knowledge of their regularity, and the forces which cause it. In the symmetry of the dividing cell the basis of that resemblance we call Heredity is contained. To imitate the morphological phenomena of life we have to devise a system which can divide. It must be able to divide, and to segment as--grossly--a vibrating plate or rod does, or as an icicle can do as it becomes ribbed in a continuous stream of water; but with this distinction, that the distribution of chemical differences and properties must simultaneously be decided and disposed in orderly relation to the pattern of the segmentation. Even if a model which would do this could be constructed it might prove to be a useful beginning.
This may be looking too far ahead. If we had to choose some one piece of more proximate knowledge which we would more especially like to acquire, Isuppose we should ask for the secret of interracial sterility. Nothing has yet been discovered to remove the grave difficulty, by which Huxley in particular was so much oppressed, that among the many varieties produced under domestication--which we all regard as analogous to the species seen in nature--no clear case of interracial sterility has been demonstrated.
The phenomenon is probably the only one to which the domesticated products seem to afford no parallel. No solution of the difficulty can be offered which has positive value, but it is perhaps worth considering the facts in the light of modern ideas. It should be observed that we are not discussing incompatibility of two species to produce offspring (a totally distinct phenomenon), but the sterility of the offspring which many of them do produce.